| Characteristics | GUHS97 |
| Gram stain | + |
| Voges proskauer tests | + |
| Methyl red tests | + |
| Citrate | + |
| Hydrogen Peroxide | + |
| Starch hydrolysis | + |
| Nitrite reductase | + |
| Nitrite reductase (NO-forming) | − |
| Alpha-L-rhamnose | + |
| Mannitol | + |