| Phosphorylation | Site | |
| TGF-β Signalling pathway | TGF-βRI | S422, S423 and S425 [54] [55] |
| MAPK Signalling pathway | ERK | T179, S204, S213 [55] S207, S203 and t178 [56] |
| P38 | S204, S208 and S213 [55] | |
| JNK | S208 and S213 [55] | |
| Cyclin-dependent kinase (CDK) family | CDK4 and CDK2 | T8, T178 and S212 [56] [57] |
| CDK8/9 | T179, S208 and S213 [57] | |
| Glycogen synthase kinase-3β (GSK-3β) | GSK-3β | T66 and T179 [55] [57] |
| Cyclin guanosine monophosphate/Protein Kinase G | cGMP/PKG | S309 and T368 [58] |